Which layer of microsporangium is nutritive in function?

The correct answer is Option (4) → Tapetum

A microsporangium of the anther has four wall layers, from outside in: epidermis, endothecium, middle layers and tapetum.

Option (4) Tapetum — Correct. The innermost layer, the tapetum, has cells with dense cytoplasm and often more than one nucleus. It nourishes the developing pollen grains.

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NCERT Extract:
Stamen, Microsporangium and Pollen Grain – “… Structure of microsporangium: In a transverse of an anther section, a typical microsporangium appears near circular in outline. It is generally surrounded by four wall layers (Figure 1.3b)– the epidermis, endothecium, middle layers and the tapetum. The outer three wall layers perform the function of protection and help in dehiscence of anther to release the pollen. The innermost wall layer is the tapetum. It nourishes the developing pollen grains.”